Tracey Lund claimed first prize at the World Nature Photography Awards with this image of gannets fishing off Scotland's Shetland Islands. Scroll through the gallery to see more winners from the competition. Limited digital manipulation was allowed.
Tracey Lund/World Nature Photography Awards
Now in its fourth year, the awards received thousands of submissions. This photo, of a yellow-billed oxpecker sitting on an African water buffalo, took silver prize in the bird behavior category.
Thomas Vijayan won silver in that category, capturing a Canadian lynx pouncing in the snow.
Thomas Vijayan/World Nature Photography Awards
This photo of a turtle foraging off San Cristóbal, one of Ecuador's Galapagos Islands, won bronze in the animals in their habitat category.
Tom Shlesinger/World Nature Photography Awards
Also shot in the Galapagos Islands, John Seager, from the UK, took gold in the amphibians and reptiles behavior category with this image, showing a lava lizard standing on a marine iguana.
John Seager/World Nature Photography Awards
Another image from the Galapagos Islands shows Sally lightfoot crabs. It won Bill Klipp the invertebrates behavior category.
Bill Klipp/World Nature Photography Awards
This photo of a leopard by a pond in Kenya's East Rift Valley won Richard Li the black and white category.
Richard Li/World Nature Photography Awards
Celia Kujala won the nature photojournalism category with this photo of a sea lion pup playing with garbage off the Coronado Islands, Baja California, Mexico.
Celia Kujala/World Nature Photography Awards
Andy Schmid, from Switzerland, took gold in the underwater category with this image of a female orca chasing herring, in Skjervøy, Norway.
Andy Schmid
An unusual plant called Thismia thaithongiana, photographed in Umphang Wildlife Sanctuary, Tak Province, Thailand, was the subject of the photo that won the plants and fungi category.
